Item 1.01 Entry into a Material Definitive Agreement.

On January 6, 2023, Keros Therapeutics, Inc. (the "Company") entered into a first amendment (the "Lease Amendment") to that certain indenture of lease, dated September 7, 2021 (as amended, the "Lease") with Revolution Labs Owner, LLC, a Delaware limited liability company (the "Landlord"), with respect to the Company's office and laboratory space consisting of approximately 35,662 square feet located at 1050 Waltham Street, Lexington, Massachusetts (the "Premises").

Under the terms of the Lease Amendment, the Company has agreed to the phased delivery of the Premises to the Company by the Landlord as follows: (i) the Landlord will deliver approximately 31,991 square feet of the Premises (the "Phase A Premises") on or before January 4, 2023 (the "Phase A Premises Commencement Date"), and (ii) the Landlord will deliver the additional approximately 3,671 rentable square feet of vivarium space (the "Phase B Premises") when the Landlord substantially completes work on the Phase B Premises, currently estimated to occur on or before May 1, 2023 (the "Phase B Premises Commencement Date").

Per the Lease Amendment, the term of the Lease will now commence on the Phase A Premises Commencement Date. The Lease has a term of eight years, measured from the Phase A Premises Commencement Date.

The Company's obligation for the payment of base rent for the Phase A Premises begins on the date which is earlier to occur of (i) eight months following the Phase B Premises Commencement Date or (ii) 11 months following the Phase A Premises Commencement Date (such earlier date, the "Phase A Rent Commencement Date"). The Company's obligation for the payment of base rent for the Phase B Premises begins on the date which is eight months following the Phase B Premises Commencement Date (the "Phase B Rent Commencement Date"). Base rent will initially be fixed at $202,084.66 per month, which will increase by approximately 3% per annum. From and after the Phase A Rent Commencement Date and until the occurrence of the Phase B Rent Commencement Date, the Company will be obligated to pay base rent at the per square foot rent for the Phase A Premises only. Upon the Phase B Rent Commencement Date, the Company will be obligated to pay the base rent for the entirety of the Premises. The Company will be obligated to pay the Landlord for certain costs, taxes and operating expenses as specified in the Lease.

The description of the Lease Amendment contained in this Current Report on Form 8-K does not purport to be complete and is qualified in its entirety by reference to the Lease Amendment, a copy of which will be filed as an exhibit to the Company's Quarterly Report on Form 10-Q for the period ending March 31, 2023.

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